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Integrative Taxonomy


Admission requirements

Advice: BSc-course Biodiversity, theory and practice of systematics or equivalent knowledge of phylogenetic reconstruction.


Coordinator: Dr. P.H. Hovenkamp


The main aim of this course is to introduce a number of approaches and methods that are currently used in both actual and historical analyses of biodiversity and evolution. The following topics will be introduced and practiced:

  • Phylogenetic analysis: concepts and applications.

  • Analysis of geographic structure and gene flow in and between populations.

  • Analysis of historic and stratigraphic patterns, paleobiogeography and –ecology.

  • Spatial pattern analyses and historical biogeography: integrative approaches to map and analyze present distributions and reconstruct past distributions.

Learning goals

Course objectives:
To provide a theoretical basis and some practical experience for the a successful MSc research project at Naturalis: good understanding of concepts of time and space; of collection data, data basing and data mining; of the practice of phylogeny reconstruction; of biogeographic methods and practice; of the integration of paleontological and actual data.

Final qualifications:
Theoretical understanding and hands on experience with current methods for phylogeny reconstruction, historical biogeography, diversity analysis in both actual and stratigraphic contexts, phylogeographical analysis, dating techniques, etc.


From 27 November 2017 to 22 December 2017. Details will be provided in due time on Blackboard; general outline is 1 week phylogeny, 1 week biogeography, 1 week speciation and population level-analysis, 1 week geological time, all with focus on current methods and concepts.

Mode of instruction

Lectures, computer exercises, literature study, demonstrations.

Assessment method

Written reports, oral presentation, and examination.


All information of lectures and papers will be available on Blackboard.

Reading list

Relevant literature will be made available on Blackboard.


Via USIS and enroll in Blackboard

Exchange and Study Abroad students, please see the Prospective students website for more information on how to apply.